The biological association that contributes directly to succession in a community is

The biological association that contributes directly to succession in a community is "competition". Competition occurs when organisms of the same or different species compete for the same resources such as food, space, or mates. In a community, when one species outcompetes another for these resources, it can lead to a shift in the balance of the community and ultimately contribute to succession. For example, if one species of plant is more efficient at using resources like sunlight and nutrients, it may outcompete other plants and become dominant in the community, altering the composition of the community over time. Therefore, competition is an important factor that drives succession in a community.